Dr Mark Stephen Harvey
Dr Mark Stephen Harvey
Department of Terrestrial Invertebrates
Western Australian Museum
Locked Bag 49, Welshpool DC
Western Australia 6986
Australia
Education
- Bachelor of Science, 1979, Monash University
- Doctor of Philosophy, 1983, Monash University [Contributions to the systematics of the Pseudoscorpionida (Arachnida): the genus Synsphyronus Chamberlin (Garypidae) and the family Sternophoridae]
Research Interests
For nearly 25 years I have maintained a research programme devoted to the systematics and biogeography of Australasian arachnids, with major contributions to the study of pseudoscorpions, spiders and water mites, as well as some of the smaller orders such as the Schizomida and Amblypygi.
I have developed an online catalog of the Pseudoscorpions of the World.
Publications
(For copyright reasons, pdf-files of publications are not available for direct download. Please send an email to Mark if you are interested in a particular publication.)
- 156. Harvey, M.S. and Leng, M.C. (2008). The first troglomorphic pseudoscorpion of the family Olpiidae (Pseudoscorpiones), with remarks on the composition of the family. Records of the Western Australian Museum 24: 379-386.155. Harvey, M.S. and Leng, M.C. (2008). Further observations on Ideoblothrus (Pseudoscorpiones: Syarinidae) from subterranean environments in Australia. Records of the Western Australian Museum 24: 379-386.154. Ott, R. and Harvey, M.S. (2008). A new species of Xestaspis (Araneae: Oonopidae) from the arid zone of Western Australia. Records of the Western Australian Museum 24: 337-342.153. Ott, R. and Harvey, M.S. (2008). A new species of Pelicinus from the Pilbara region of Western Australia (Araneae: Oonopidae). Arthropoda Selecta 17: 81-85.152. Rahmadi, C. and Harvey, M.S. (2008). A first epigean species of Stygophrynus Kraepelin (Amblypygi: Charontidae) from Java and adjacent islands, Indonesia, with notes on S. dammermani Roewer, 1928. Raffles Bulletin of Zoology 56: 281-288.151. Murienne, J., Harvey, M.S. and Giribet, G. (2008). First molecular phylogeny of the major clades of Pseudoscorpiones (Arthropoda: Chelicerata). Molecular Phylogenetics and Evolution 49: 170-184.150. Moir, M.L. and Harvey, M.S. (2008). Discovery of the pill millipede genus Epicyliosoma (Diplopoda: Sphaerotheriida: Sphaerotheriidae) in Western Australia, with the description of a new species. Records of the Western Australian Museum 24: 113-119.149. Edward, K.L. and Harvey, M.S. (2008). Short-range endemism in hypogean environments: the pseudoscorpion genera Tyrannochthonius and Lagynochthonius (Pseudoscorpiones: Chthoniidae) in the semi-arid zone of Western Australia. Invertebrate Systematics 22: 259-293.148. Barranco, P. and Harvey, M.S. (2008). The first indigenous species of Eukoenenia (Palpigradi: Eukoeneniidae) from Australia. Invertebrate Systematics 22: 227-233.147. Harvey, M.S., Berry, O., Edward, K.L. and Humphreys, G. (2008). Molecular and morphological systematics of hypogean schizomids (Schizomida : Hubbardiidae) in semi-arid Australia. Invertebrate Systematics 22: 167-194.146. Boulton, A.J., Fenwick, G., Hancock, P. J. and Harvey, M.S. (2008). Biodiversity, functional roles and ecosystem services of groundwater invertebrates. Invertebrate Systematics 22: 106-116.145. Rix, M.G., Harvey, M.S. and Roberts, J.D. (2008). Molecular phylogenetics of the spider family Micropholcommatidae (Arachnida: Araneae) using nuclear rRNA genes (18S and 28S). Molecular Phylogenetics and Evolution 46: 1031-1048.
2007
144. Harvey, M.S. and Edward, K.L. (2007). Three new species of cavernicolous goblin spiders (Araneae, Oonopidae) from Australia. Records of the Western Australian Museum 24: 9-17.143. Harvey, M.S. (2007). The smaller arachnid orders: diversity, descriptions and distributions from Linnaeus to the present (1758 to 2007). Zootaxa 1668: 363-380.142. Harvey, M.S., Austin, A.D. and Adams, M. (2007). The systematics and biology of the spider genus Nephila (Araneae: Nephilidae) in the Australasian region. Invertebrate Systematics 21: 407-451.141. Harvey, M.S. and Volschenk, E.S. (2007). The systematics of the Gondwanan pseudoscorpion family Hyidae (Pseudoscorpiones: Neobisioidea): new data and a revised phylogenetic hypothesis. Invertebrate Systematics 21: 365-406.140. Harvey, M.S. and Volschenk, E.S. (2007). A review of some Australasian Chernetidae: Sundochernes, Troglochernes and a new genus (Pseudoscorpiones). Journal of Arachnology 35: 238-277.139. Fontaine, B., Bouchet, P., van Achterberg, K., Alonso-Zarazaga, M.A., Araujo, R.A., Aspöck, U., Audisio, P., Aukema, B., Bailly, N., Balsamo, M., Bank, R.A., Barnard, P., Belfiore, C., Bogdanowicz, W., Bongers, T., Boxshall, G., Burckhardt, D., Camicas, J.-L., Chylarecki, P., Crucitti, P., Deharveng, L., Dubois, A., Enghoff, H., Faubel, A., Fochetti, R., Gargominy, O., Gibson, D., Gibson, R., Gómez López, M.S., Goujet, D., Harvey, M.S., Heller, K.-G., van Helsdingen, P., Hoch, H., de Jong, H., de Jong, Y., Karsholt, O., Los, W., Lundqvist, L., Magowski, W., Manconi, R., Martens, J., Massard, J.A., Massard-Geimer, G., McInnes, S.J., Mendes, L.F., Mey, E., Michelsen, V., Minelli, A., Nielsen, C., Nieto Nafría, J.M., van Nieukerken, E., Noyes, J., Pape, T., Pohl, H., De Prins, W., Ricci, C., Roselaar, C., Rota, E., Schmidt-Rhaesa, A., Segers, H., zur Strassen, R., Szeptycki, A., Thibaud, J.-M., Thomas, A., Timm, T., van Tol, J., Vervoort, W. and Willmann, R. (2007). The European Union’s 2010 target: putting rare species in focus. Biological Conservation 139: 167-185.138. Harvey, M.S., Waldock, J.M., Teale, R.J. and Webber, J. (2007). New distribution records of the intertidal pseudoscorpion Parahya submersa (Pseudoscorpiones: Parahyidae). Records of the Western Australian Museum 23: 393-395.137. Rahmadi, C. and Harvey, M.S. (2007). The female of Phrynus exsul (Amblypygi, Phrynidae) from Indonesia. Journal of Arachnology 35: 137-142.136. Moulds, T.A., Murphy, N., Adams, M., Reardon, T., Harvey, M.S., Jennings, J. and Austin, A.D. (2007). Phylogeography of cave pseudoscorpions in southern Australia. Journal of Biogeography 34: 951-962.135. Harvey, M.S. and Edward, K.L. (2007). A review of the pseudoscorpion genus Ideoblothrus (Pseudoscorpiones, Syarinidae) from western and northern Australia. Journal of Natural History 41: 445-472.134. Harvey, M.S., Barba D., R., Muchmore, W.B. and Perez G., A. (2007). Pseudalbiorix, a new genus of Ideoroncidae (Pseudoscorpiones, Neobisioidea) from central America. Journal of Arachnology 34: 610-626.2006
133. Dippenaar-Schoeman, A., González Reyes, A.P. and Harvey, M.S. (2006). A check-list of the Solifugae (sun-spiders) of South Africa (Arachnida: Solifugae). African Plant Protection 12: 70-92.132. Harvey, M.S. and Mahnert, V. (2006). The systematic position of the Amazonian species of Albiorix (Pseudoscorpiones, Ideoroncidae). Journal of Arachnology 34: 227-230.131. Harvey, M.S. and Mould, L.G. (2006). A new troglomorphic species of Austrochthonius (Pseudoscorpiones: Chthoniidae) from Australia, with remarks on Chthonius caecus. Records of the Western Australian Museum 23: 205-211.130. Harvey, M.S., Stahlavsky, F. and Theron, P.D. (2006). The distribution of Eukoenenia mirabilis (Palpigradi: Eukoeneniidae): a widespread tramp. Records of the Western Australian Museum 23: 199-203.129. Harvey, M.S. (2006). New species and records of the pseudoscorpion family Menthidae (Pseudoscorpiones). Records of the Western Australian Museum 23: 167-174.128. Stahlavsky, F., Kral, J., Harvey, M.S. and Haddad, C.R. (2006). A karyotype study on the pseudoscorpion families Geogarypidae, Garypinidae and Olpiidae (Arachnida: Pseudoscorpiones). European Journal of Entomology 103: 277-289.127. Zaragoza, J. and Harvey, M.S. (2006). The first record of the genus Mundochthonius Chamberlin (Pseudoscorpiones: Chthoniidae) from Spain: Mundochthonius gallaecicus sp. nov. Revista Iberica de Aracnologia 12: 17-23.126. Harvey, M.S. (2006). A new species of Lechytia from eastern Australia (Pseudoscorpiones: Lechytiidae). Records of the Western Australian Museum 23: 13-18.125. Harvey, M.S. (2006). The schizomid fauna of the Arabian Peninsula and Somalia (Arachnida: Schizomida: Hubbardiidae). Fauna of Arabia 21: 167-177.124. Murphy, N.P., Framenau, V.W., Donnellan, S.C., Harvey, M.S., Park, Y.-C., Austin, A.D. (2006). Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12S rRNA, 28S rRNA and NADH1 genes: implications for classification, biogeography and the evolution of web building behaviour. Molecular Phylogeny and Evolution 38: 583-602.2004
123. Harvey, M.S., Waldock, J.M., Guthrie, N.A., Durrant, B.J. and McKenzie, N.L. (2004). Patterns in the composition of ground-dwelling araneomorph spider communities in the Western Australian wheatbelt. Records of the Western Australian Museum, Supplement 67: 257-291.122. Harvey, M.S. (2004). Remarks on the New World pseudoscorpion genera Parawithius and Victorwithius, with a new genus bearing a remarkable sternal modification (Pseudoscorpiones, Withiidae). Journal of Arachnology 32: 436-456.121. Boulton, A.J., Harvey, M.S. and Proctor, H. (2004). Of spates and species: responses by interstitial water mites to simulated spates in a subtropical Australian river. Experimental and Applied Acarology 34: 149-169.120. Coddington, J.A., Giribet, G., Harvey, M.S., Prendini, L. and Walter, D.E. (2004). Arachnida. In: Cracraft, J. and Donoghue, M.J. (eds), Assembling the Tree of Life. Oxford University Press, New York.119. Yeates, D.K., Harvey, M.S. and Austin, A.D. (2004). New estimates for terrestrial arthropod species-richness in Australia. Records of the South Australian Museum Monograph Series 7: 231-241.118. Balit, C.R., Harvey, M.S., Waldock, J.M. and Isbister, G.K. (2004). Prospective study of centipede bites in Australia. Journal of Toxicology - Clinical Toxicology 42: 41-48.2003
117. Harvey, M.S. (2003). Davecookia, a new genus of the water mite family Athienemanniidae (Acarina: Hydracarina) from Australia. In: Smith, I.M. (ed.), An acarological tribute to David R. Cook - from Yankee Springs to Wheeny Creek: 151-153. Indira Publishing House: West Bloomfield.116. Boulton, A.J. and Harvey, M.S. (2003). Effects of a simulated spate on water mites in the hyporheic zone of an Australian subtropical river. In: Smith, I.M. (ed.), An acarological tribute to David R. Cook - from Yankee Springs to Wheeny Creek: 57-73. Indira Publishing House: West Bloomfield.115. Harvey, M.S. (2003). Rhagodes Pocock, 1897 (Solifugae: Rhagodidae), proposed conservation through the suppression of Rhax Hermann, 1804. Bulletin of Zoological Nomenclature 60: 117-118.114. Isbister, G.K., Volschenk, E.S., Balit, C.R. and Harvey, M.S. (2003). Australian scorpion stings: a prospective study of definite stings. Toxicon 41: 877-883.112. Harvey, M.S. (2003). Zeriassa Pocock, 1897 (Solifugae: Solpugidae), proposed conservation through the suppression of Canentis Pavesi, 1897. Bulletin of Zoological Nomenclature 60: 26-27.2002
111. Harvey, M.S. (2002). The first Old World species of Phrynidae (Amblypygi): Phrynus exsul sp. nov. from Indonesia. Journal of Arachnology 30: 470-474.109. Harvey, M.S. (2002). The neglected cousins: what do we know about the smaller arachnid orders? Journal of Arachnology 30: 357-372.108. Harvey, M.S. (2002). Short-range endemism amongst the Australian fauna: examples from non-marine environments. Invertebrate Systematics 16: 555-570.107. Austin, A.D., Fisher, R.H., Harvey, M.S., Hirst, D.B., Locket, N.A., Madden, C.P., Reay, F. and Zeidler, W. (2002). Terrestrial and freshwater invertebrates. In: Davies, M., Twidale, C.R. and Tyler, M.J. (eds), Natural History of the Kangaroo Island, 2nd edition: 123-141. Royal Society of South Australia: Adelaide.106. Harvey, M.S. and Volschenk, E.S. (2002). A forgotten scorpion: the identity of Buthus flavicruris Rainbow, 1896 (Scorpiones), with notes on Urodacus manicatus. Records of the Western Australian Museum 21: 105-106.105. Harvey, M.S. (2002). A new species of Austrarchaea (Araneae: Archaeidae) from Western Australia. Records of the Western Australian Museum 21: 35-37.104. Harvey, M.S. (2002). Nomenclatural notes on Solifugae, Amblypygi, Uropygi and Araneae (Arachnida). Records of the Western Australian Museum 20: 449-459.103. Main, B.Y., Harvey, M.S. and Waldock, J.M. (2002). The distribution of the Western Australian pill millipede, Cynotelopus notabilis Jeekel (Sphaerotheriidae). Records of the Western Australian Museum 20: 383-385.2001
102. Harvey, M.S. (2001). Notes on the spider genus Symphytognatha (Araneae: Symphytognathidae) in Western Australia. Records of the Western Australian Museum 20: 345-347.101. Harvey, M.S. (2001). New cave-dwelling schizomids (Schizomida: Hubbardiidae) from Australia. Records of the Western Australian Museum, Supplement 64: 171-185.100. Harvey, M.S. (2001). The Schizomida (Arachnida) of the Seychelle Islands. Invertebrate Taxonomy 15: 681-693.99. Harvey, M.S. (2001). Ammotrecha Banks, 1900 and Ammotrechula Roewer, 1934 (Arachnida, Solifugae): proposed conservation by the designation of Galeodes limbata Lucas, 1835 as the type species of Ammotrecha; and Eremobates Banks, 1900 and Eremorhax Roewer, 1934: proposed conservation by the designation of Galeodes pallipes Say, 1823 as the type species of Eremobates. Bulletin of Zoological Nomenclature 58: 196-201.98. Derraik, J.G.B., Barratt, B.I.P., Sirvid, P., MacFarlane, R.P., Patrick, B.H., Early, J., Eyles, A.C., Johns, P.M., Fraser, P.M., Barker, G.M., Henderson, R., Dale, P.J., Harvey, M.S., Fenwick, G., McLellan, I.D., Dickinson, K.J.M., Closs, G.P. (2001). Invertebrate survey of a modified native shrubland, Brookdale Covenant, Rock & Pillar Range, Otago, New Zealand. New Zealand Journal of Zoology 28: 273-290.97. Volschenk, E.S., Locket, N.A. and Harvey, M.S. (2001). Liocheles polisorum n. sp., first record of a troglobitic ischnurid scorpion from Australasia (Scorpiones: Ischnuridae). In: Fet, V. and Selden, P.A. (eds), Scorpions 2001. In memorium Gary A. Polis: 161-170. British Arachnological Society: Burnham Beeches, Bucks.2000
96. Harvey, M.S. (2000). Brignolizomus and Attenuizomus, new schizomid genera from Australia (Schizomida Hubbardiidae). Memorie della Societa Entomologica Italiana, Supplement 78: 329-338.95. Harvey, M.S. (2000). A review of the Australian schizomid genus Notozomus (Hubbardiidae). Memoirs of the Queensland Museum 46: 161-174.94. Harvey, M.S., Sampey, A., West, P.L.J. and Waldock, J.M. (2000). The Chilopoda and Diplopoda of the southern Carnarvon Basin, Western Australia. Records of the Western Australian Museum, Supplement 61: 323-333.93. Harvey, M.S., Sampey, A., West, P.L.J. and Waldock, J.M. (2000). Araneomorph spiders from the southern Carnarvon Basin, Western Australia: a consideration of regional biogeographic relationships. Records of the Western Australian Museum, Supplement 61: 295-321.92. Halse, S.A., Shiel, R.J., Storey, A.W., Edward, D.H.D., Lansbury, I., Cale, D.J. and Harvey, M.S. (2000). Aquatic invertebrates and waterbirds of wetlands and rivers of the southern Carnarvon Basin, Western Australia. Records of the Western Australian Museum, Supplement 61: 217-265.91. Burbidge, A.H., McKenzie, N.L. and Harvey, M.S. (2000). A biogeographic survey of the southern Carnarvon Basin, Western Australia: background and methods. Records of the Western Australian Museum, Supplement 61: 1-12.90. Dippenaar-Schoeman, A.S. and Harvey, M.S. (2000). A check list of the pseudoscorpions of South Africa (Arachnida: Pseudoscorpiones). Koedoe 43: 89-102.89. Harvey, M.S. (2000). From Siam to Rapa Nui - the identity and distribution of Geogarypus longidigitatus (Rainbow) (Pseudoscorpiones: Geogarypidae). Bulletin of the British Arachnological Society 11: 377-384.88. Harvey, M.S., Shear, W.S. and Hoch, H. (2000). Onychophora, Arachnida, Myriapods and Insecta. In: Wilkens, H., Culver, D.C. and Humphreys, W.F. (eds), Subterranean Ecosystems, vol. 30: 79-94. Elsevier: Amsterdam.87. Adis, J. and Harvey, M.S. (2000). How many Arachnida and Myriapoda are there world-wide and in Amazonia? Studies on Neotropical Fauna and Environment 35: 139-141.86. Volschenk, E.S., Smith, G.T. and Harvey, M.S. (2000). A new species of Urodacus from Western Australia, with descriptive notes on Urodacus megamastigus (Scorpiones: Urodacidae). Records of the Western Australian Museum 20: 57-68.85. Cook, D.R., Smith, I.M. and Harvey, M.S. (2000). Assessment of lateral compression of the idiosoma in adult water mites as a taxonomic character and reclassification of Frontipodopsis Walter, Wettina Piersig and some other basal Hygrobatoidea (Acari : Hydrachnida). Invertebrate Taxonomy 14: 433-448.84. Harvey, M.S. and Waldock, J.M. (2000). A review of the spider genus Yoroa Baert (Araneae: Theridiidae: Hadrotarsinae). Australian Journal of Entomology 39: 58-61.1999
83. Harvey, M.S. (1999). The Asian species of Microcreagris Balzan (Pseudoscorpiones: Neobisiidae) described by J.C. Chamberlin. Acta Arachnologica 48: 93-105.82. Harvey, M.S. (1999). The identity of the Indonesian pseudoscorpions Lagynochthonius johni (Redikorzev) and Lagynochthonius roeweri Chamberlin (Pseudoscorpiones: Chthoniidae). Bulletin of the British Arachnological Society 11: 247-250.1998
81. Harvey, M.S. (1998). Pseudoscorpion groups with bipolar distributions: a new genus from Tasmania related to the Holarctic Syarinus (Arachnida, Pseudoscorpiones, Syarinidae). Journal of Arachnology 26: 429-441.80. Harvey, M.S. and West, P.J. (1998). New species of Charon (Amblypygi: Charontidae) from northern Australia and Christmas Island. Journal of Arachnology 26: 273-284.78. Harvey, M.S. (1998). A review of the Australasian species of Anapistula Gertsch (Araneae: Symphytognathidae). Records of the Western Australian Museum 19: 111-120.77. Harvey, M.S. (1998). Unusual new water mites (Acari: Hydracarina) from Australia, Part 1. Records of the Western Australian Museum 19: 91-106.76. Austin, A.D., Gibson, G.A.P. and Harvey, M.S. (1998). Synopsis of Australian Calymmochilus Masi (Hymenoptera: Eupelmidae), description of a new species from Western Australia associated with a pseudoscorpion, and review of pseudoscorpion parasites. Journal of Natural History 32: 329-350.1997
74. Harvey, M.S., Waldock, J.M., How, R.A., Dell, J. and Kostas, E. (1997). Biodiversity and biogeographic relationships of selected invertebrates from urban bushland remnants, Perth, Western Australia. Memoirs of the Museum of Victoria 56: 275-280.1996
73. Harvey, M.S. (1996). The first record of the Fiddle-back Spider Loxosceles rufescens (Araneae: Sicariidae) from Western Australia. Records of the Western Australian Museum 18: 223-224.72. Austin, A.D., Jennings, J.T. and Harvey, M.S. (1996). Pseudofoenus Kieffer, 1902 (Insecta, Hymenoptera): proposed designation of Foenus unguiculatus Westwood, 1841 as the type species. Bulletin of Zoological Nomenclature 53: 261-263.71. Austin, A.D., Baker, G.H., Colman, P.H., Fisher, R.H., Harvey, M.S., Hirst, D.B., Locket, N.A. and Reay, F. (1996). Terrestrial invertebrates. In: Natural History of the Flinders Ranges: 113-126. Royal Society of South Australia: Adelaide.70. Harvey, M.S. (1996). Small arachnids and their value in Gondwanan biogeographic studies. In: Hopper, S.D., Chappill, J.A., Harvey, M.S. and George, A.S. (eds), Gondwanan Heritage: 155-162. Surrey Beatty and Sons: Sydney.