Dr Mark Stephen Harvey
Senior Curator
Department of Terrestrial Invertebrates
Western Australian Museum
Locked Bag 49, Welshpool DC
Western Australia 6986
Australia
Education
- Bachelor of Science, 1979, Monash University
- Doctor of Philosophy, 1983, Monash University [Contributions to the systematics of the Pseudoscorpionida (Arachnida): the genus Synsphyronus Chamberlin (Garypidae) and the family Sternophoridae]
Research Interests
For nearly 25 years I have maintained a research programme devoted to the systematics and biogeography of Australasian arachnids, with major contributions to the study of pseudoscorpions, spiders and water mites, as well as some of the smaller orders such as the Schizomida and Amblypygi.
Publications
(For copyright reasons, pdf-files of publications are not available for direct download. Please send an email to Mark if you are interested in a particular publication.)
- 132. Harvey, M.S. and Mahnert, V. (2006). The systematic position of the Amazonian species of Albiorix (Pseudoscorpiones, Ideoroncidae). Journal of Arachnology 34: 227-230.131. Harvey, M.S. and Mould, L.G. (2006). A new troglomorphic species of Austrochthonius (Pseudoscorpiones: Chthoniidae) from Australia, with remarks on Chthonius caecus. Records of the Western Australian Museum 23: 205-211.130. Harvey, M.S., Stahlavsky, F. and Theron, P.D. (2006). The distribution of Eukoenenia mirabilis (Palpigradi: Eukoeneniidae): a widespread tramp. Records of the Western Australian Museum 23: 199-203.129. Harvey, M.S. (2006). New species and records of the pseudoscorpion family Menthidae (Pseudoscorpiones). Records of the Western Australian Museum 23: 167-174.128. Stahlavsky, F., Kral, J., Harvey, M.S. and Haddad, C.R. (2006). A karyotype study on the pseudoscorpion families Geogarypidae, Garypinidae and Olpiidae (Arachnida: Pseudoscorpiones). European Journal of Entomology 103: 277-289.127. Zaragoza, J. and Harvey, M.S. (2006). The first record of the genus Mundochthonius Chamberlin (Pseudoscorpiones: Chthoniidae) from Spain: Mundochthonius gallaecicus sp. nov. Revista Iberica de Aracnologia 12: 17-23.126. Harvey, M.S. (2006). A new species of Lechytia from eastern Australia (Pseudoscorpiones: Lechytiidae). Records of the Western Australian Museum 23: 13-18.125. Harvey, M.S. (2006). The schizomid fauna of the Arabian Peninsula and Somalia (Arachnida: Schizomida: Hubbardiidae). Fauna of Arabia 21: 167-177.124. Murphy, N.P., Framenau, V.W., Donnellan, S.C., Harvey, M.S., Park, Y.-C., Austin, A.D. (2006). Phylogenetic reconstruction of the wolf spiders (Araneae: Lycosidae) using sequences from the 12S rRNA, 28S rRNA and NADH1 genes: implications for classification, biogeography and the evolution of web building behaviour. Molecular Phylogeny and Evolution 38: 583-602.
2004
123. Harvey, M.S., Waldock, J.M., Guthrie, N.A., Durrant, B.J. and McKenzie, N.L. (2004). Patterns in the composition of ground-dwelling araneomorph spider communities in the Western Australian wheatbelt. Records of the Western Australian Museum, Supplement 67: 257-291.122. Harvey, M.S. (2004). Remarks on the New World pseudoscorpion genera Parawithius and Victorwithius, with a new genus bearing a remarkable sternal modification (Pseudoscorpiones, Withiidae). Journal of Arachnology 32: 436-456.121. Boulton, A.J., Harvey, M.S. and Proctor, H. (2004). Of spates and species: responses by interstitial water mites to simulated spates in a subtropical Australian river. Experimental and Applied Acarology 34: 149-169.120. Coddington, J.A., Giribet, G., Harvey, M.S., Prendini, L. and Walter, D.E. (2004). Arachnida. In: Cracraft, J. and Donoghue, M.J. (eds), Assembling the Tree of Life. Oxford University Press, New York.119. Yeates, D.K., Harvey, M.S. and Austin, A.D. (2004). New estimates for terrestrial arthropod species-richness in Australia. Records of the South Australian Museum Monograph Series 7: 231-241.118. Balit, C.R., Harvey, M.S., Waldock, J.M. and Isbister, G.K. (2004). Prospective study of centipede bites in Australia. Journal of Toxicology - Clinical Toxicology 42: 41-48.2003
117. Harvey, M.S. (2003). Davecookia, a new genus of the water mite family Athienemanniidae (Acarina: Hydracarina) from Australia. In: Smith, I.M. (ed.), An acarological tribute to David R. Cook - from Yankee Springs to Wheeny Creek: 151-153. Indira Publishing House: West Bloomfield.116. Boulton, A.J. and Harvey, M.S. (2003). Effects of a simulated spate on water mites in the hyporheic zone of an Australian subtropical river. In: Smith, I.M. (ed.), An acarological tribute to David R. Cook - from Yankee Springs to Wheeny Creek: 57-73. Indira Publishing House: West Bloomfield.115. Harvey, M.S. (2003). Rhagodes Pocock, 1897 (Solifugae: Rhagodidae), proposed conservation through the suppression of Rhax Hermann, 1804. Bulletin of Zoological Nomenclature 60: 117-118.114. Isbister, G.K., Volschenk, E.S., Balit, C.R. and Harvey, M.S. (2003). Australian scorpion stings: a prospective study of definite stings. Toxicon 41: 877-883.112. Harvey, M.S. (2003). Zeriassa Pocock, 1897 (Solifugae: Solpugidae), proposed conservation through the suppression of Canentis Pavesi, 1897. Bulletin of Zoological Nomenclature 60: 26-27.2002
111. Harvey, M.S. (2002). The first Old World species of Phrynidae (Amblypygi): Phrynus exsul sp. nov. from Indonesia. Journal of Arachnology 30: 470-474.109. Harvey, M.S. (2002). The neglected cousins: what do we know about the smaller arachnid orders? Journal of Arachnology 30: 357-372.108. Harvey, M.S. (2002). Short-range endemism amongst the Australian fauna: examples from non-marine environments. Invertebrate Systematics 16: 555-570.107. Austin, A.D., Fisher, R.H., Harvey, M.S., Hirst, D.B., Locket, N.A., Madden, C.P., Reay, F. and Zeidler, W. (2002). Terrestrial and freshwater invertebrates. In: Davies, M., Twidale, C.R. and Tyler, M.J. (eds), Natural History of the Kangaroo Island, 2nd edition: 123-141. Royal Society of South Australia: Adelaide.106. Harvey, M.S. and Volschenk, E.S. (2002). A forgotten scorpion: the identity of Buthus flavicruris Rainbow, 1896 (Scorpiones), with notes on Urodacus manicatus. Records of the Western Australian Museum 21: 105-106.105. Harvey, M.S. (2002). A new species of Austrarchaea (Araneae: Archaeidae) from Western Australia. Records of the Western Australian Museum 21: 35-37.104. Harvey, M.S. (2002). Nomenclatural notes on Solifugae, Amblypygi, Uropygi and Araneae (Arachnida). Records of the Western Australian Museum 20: 449-459.103. Main, B.Y., Harvey, M.S. and Waldock, J.M. (2002). The distribution of the Western Australian pill millipede, Cynotelopus notabilis Jeekel (Sphaerotheriidae). Records of the Western Australian Museum 20: 383-385.2001
102. Harvey, M.S. (2001). Notes on the spider genus Symphytognatha (Araneae: Symphytognathidae) in Western Australia. Records of the Western Australian Museum 20: 345-347.101. Harvey, M.S. (2001). New cave-dwelling schizomids (Schizomida: Hubbardiidae) from Australia. Records of the Western Australian Museum, Supplement 64: 171-185.100. Harvey, M.S. (2001). The Schizomida (Arachnida) of the Seychelle Islands. Invertebrate Taxonomy 15: 681-693.99. Harvey, M.S. (2001). Ammotrecha Banks, 1900 and Ammotrechula Roewer, 1934 (Arachnida, Solifugae): proposed conservation by the designation of Galeodes limbata Lucas, 1835 as the type species of Ammotrecha; and Eremobates Banks, 1900 and Eremorhax Roewer, 1934: proposed conservation by the designation of Galeodes pallipes Say, 1823 as the type species of Eremobates. Bulletin of Zoological Nomenclature 58: 196-201.98. Derraik, J.G.B., Barratt, B.I.P., Sirvid, P., MacFarlane, R.P., Patrick, B.H., Early, J., Eyles, A.C., Johns, P.M., Fraser, P.M., Barker, G.M., Henderson, R., Dale, P.J., Harvey, M.S., Fenwick, G., McLellan, I.D., Dickinson, K.J.M., Closs, G.P. (2001). Invertebrate survey of a modified native shrubland, Brookdale Covenant, Rock & Pillar Range, Otago, New Zealand. New Zealand Journal of Zoology 28: 273-290.97. Volschenk, E.S., Locket, N.A. and Harvey, M.S. (2001). Liocheles polisorum n. sp., first record of a troglobitic ischnurid scorpion from Australasia (Scorpiones: Ischnuridae). In: Fet, V. and Selden, P.A. (eds), Scorpions 2001. In memorium Gary A. Polis: 161-170. British Arachnological Society: Burnham Beeches, Bucks.2000
96. Harvey, M.S. (2000). Brignolizomus and Attenuizomus, new schizomid genera from Australia (Schizomida Hubbardiidae). Memorie della Societa Entomologica Italiana, Supplement 78: 329-338.95. Harvey, M.S. (2000). A review of the Australian schizomid genus Notozomus (Hubbardiidae). Memoirs of the Queensland Museum 46: 161-174.94. Harvey, M.S., Sampey, A., West, P.L.J. and Waldock, J.M. (2000). The Chilopoda and Diplopoda of the southern Carnarvon Basin, Western Australia. Records of the Western Australian Museum, Supplement 61: 323-333.93. Harvey, M.S., Sampey, A., West, P.L.J. and Waldock, J.M. (2000). Araneomorph spiders from the southern Carnarvon Basin, Western Australia: a consideration of regional biogeographic relationships. Records of the Western Australian Museum, Supplement 61: 295-321.92. Halse, S.A., Shiel, R.J., Storey, A.W., Edward, D.H.D., Lansbury, I., Cale, D.J. and Harvey, M.S. (2000). Aquatic invertebrates and waterbirds of wetlands and rivers of the southern Carnarvon Basin, Western Australia. Records of the Western Australian Museum, Supplement 61: 217-265.91. Burbidge, A.H., McKenzie, N.L. and Harvey, M.S. (2000). A biogeographic survey of the southern Carnarvon Basin, Western Australia: background and methods. Records of the Western Australian Museum, Supplement 61: 1-12.90. Dippenaar-Schoeman, A.S. and Harvey, M.S. (2000). A check list of the pseudoscorpions of South Africa (Arachnida: Pseudoscorpiones). Koedoe 43: 89-102.89. Harvey, M.S. (2000). From Siam to Rapa Nui - the identity and distribution of Geogarypus longidigitatus (Rainbow) (Pseudoscorpiones: Geogarypidae). Bulletin of the British Arachnological Society 11: 377-384.88. Harvey, M.S., Shear, W.S. and Hoch, H. (2000). Onychophora, Arachnida, Myriapods and Insecta. In: Wilkens, H., Culver, D.C. and Humphreys, W.F. (eds), Subterranean Ecosystems, vol. 30: 79-94. Elsevier: Amsterdam.87. Adis, J. and Harvey, M.S. (2000). How many Arachnida and Myriapoda are there world-wide and in Amazonia? Studies on Neotropical Fauna and Environment 35: 139-141.86. Volschenk, E.S., Smith, G.T. and Harvey, M.S. (2000). A new species of Urodacus from Western Australia, with descriptive notes on Urodacus megamastigus (Scorpiones: Urodacidae). Records of the Western Australian Museum 20: 57-68.85. Cook, D.R., Smith, I.M. and Harvey, M.S. (2000). Assessment of lateral compression of the idiosoma in adult water mites as a taxonomic character and reclassification of Frontipodopsis Walter, Wettina Piersig and some other basal Hygrobatoidea (Acari : Hydrachnida). Invertebrate Taxonomy 14: 433-448.84. Harvey, M.S. and Waldock, J.M. (2000). A review of the spider genus Yoroa Baert (Araneae: Theridiidae: Hadrotarsinae). Australian Journal of Entomology 39: 58-61.1999
83. Harvey, M.S. (1999). The Asian species of Microcreagris Balzan (Pseudoscorpiones: Neobisiidae) described by J.C. Chamberlin. Acta Arachnologica 48: 93-105.82. Harvey, M.S. (1999). The identity of the Indonesian pseudoscorpions Lagynochthonius johni (Redikorzev) and Lagynochthonius roeweri Chamberlin (Pseudoscorpiones: Chthoniidae). Bulletin of the British Arachnological Society 11: 247-250.1998
81. Harvey, M.S. (1998). Pseudoscorpion groups with bipolar distributions: a new genus from Tasmania related to the Holarctic Syarinus (Arachnida, Pseudoscorpiones, Syarinidae). Journal of Arachnology 26: 429-441.80. Harvey, M.S. and West, P.J. (1998). New species of Charon (Amblypygi: Charontidae) from northern Australia and Christmas Island. Journal of Arachnology 26: 273-284.78. Harvey, M.S. (1998). A review of the Australasian species of Anapistula Gertsch (Araneae: Symphytognathidae). Records of the Western Australian Museum 19: 111-120.77. Harvey, M.S. (1998). Unusual new water mites (Acari: Hydracarina) from Australia, Part 1. Records of the Western Australian Museum 19: 91-106.76. Austin, A.D., Gibson, G.A.P. and Harvey, M.S. (1998). Synopsis of Australian Calymmochilus Masi (Hymenoptera: Eupelmidae), description of a new species from Western Australia associated with a pseudoscorpion, and review of pseudoscorpion parasites. Journal of Natural History 32: 329-350.1997
74. Harvey, M.S., Waldock, J.M., How, R.A., Dell, J. and Kostas, E. (1997). Biodiversity and biogeographic relationships of selected invertebrates from urban bushland remnants, Perth, Western Australia. Memoirs of the Museum of Victoria 56: 275-280.1996
73. Harvey, M.S. (1996). The first record of the Fiddle-back Spider Loxosceles rufescens (Araneae: Sicariidae) from Western Australia. Records of the Western Australian Museum 18: 223-224.72. Austin, A.D., Jennings, J.T. and Harvey, M.S. (1996). Pseudofoenus Kieffer, 1902 (Insecta, Hymenoptera): proposed designation of Foenus unguiculatus Westwood, 1841 as the type species. Bulletin of Zoological Nomenclature 53: 261-263.71. Austin, A.D., Baker, G.H., Colman, P.H., Fisher, R.H., Harvey, M.S., Hirst, D.B., Locket, N.A. and Reay, F. (1996). Terrestrial invertebrates. In: Natural History of the Flinders Ranges: 113-126. Royal Society of South Australia: Adelaide.70. Harvey, M.S. (1996). Small arachnids and their value in Gondwanan biogeographic studies. In: Hopper, S.D., Chappill, J.A., Harvey, M.S. and George, A.S. (eds), Gondwanan Heritage: 155-162. Surrey Beatty and Sons: Sydney.69. Hopper, S.D., Harvey, M.S., Chappill, J.A., Main, A.R. and Main, B.Y. (1996). The Western Australian biota as Gondwanan heritage. In: Hopper, S.D., Chappill, J.A., Harvey, M.S. and George, A.S. (eds), Gondwanan Heritage: 1-46. Surrey Beatty and Sons: Sydney.68. Harvey, M.S. (1996). The biogeography of Gondwanan pseudoscorpions. Revue Suisse de Zoologie, hors serie 1: 255-264.67. Harvey, M.S. (1996). Two new species of the water mite genus Penemideopsis from Western Australia (Acarina: Mideopsidae). Records of the Western Australian Museum 17: 443-446.66. Harvey, M.S. (1996). A review of the water mite family Pionidae in Australia (Acarina: Hygrobatoidea). Records of the Western Australian Museum 17: 361-393.65. Harvey, M.S. (1996). A revised systematic placement for Austrotrombella Southcott (Acarina: Hydryphantidae). Transactions of the Royal Society of South Australia 120: 37-40.1995
64. Cole, D.C., Elgar, M.A. and Harvey, M.S. (1995). Ecological associations between Australian pseudoscorpions and ants. Psyche 101: 221-227.63. Harvey, M.S. and Selden, P.A. (1995). Nyranytarbus, replacement name for Hemiphrynus Fric, 1901 (Trigonotarbida: Eophrynidae). Bulletin of the British Arachnological Society 10: 74.62. Harvey, M.S. (1995). The systematics of the spider family Nicodamidae (Araneae: Amaurobioidea). Invertebrate Taxonomy 9: 279-386.61. Harvey, M.S. (1995). Barbaraella, gen. nov. and Cacoxylus Beier (Pseudoscorpionida: Chernetidae), two remarkable sexually dimorphic pseudoscorpions from Australasia. Records of the Western Australian Museum, Supplement 52: 199-208.60. Harvey, M.S. and Humphreys, W.F. (1995). Notes on the genus Draculoides Harvey (Schizomida: Hubbardiidae), with the description of a new troglobitic species. Records of the Western Australian Museum, Supplement 52: 183-189.1994
59. Harvey, M.S. (1994). Rediscovered pseudoscorpion type material described by Beier from southeastern Queensland (Arachnida: Pseudoscorpionida). Memoirs of the Queensland Museum 37: 155-156.58. Harvey, M.S. (1994). Redescription and the systematic position of the Brazilian genus Xenochernes Feio (Pseudoscorpionida: Chernetidae). Journal of Arachnology 22: 131-137.1993
57. Harvey, M.S., Gray, M.R., Hunt, G.S. and Lee, D.C. (1993). The cavernicolous Arachnida and Myriapoda of Cape Range, Western Australia. Records of the Western Australian Museum, Supplement 45: 129?144.56. Harvey, M.S. and Parnaby, H.E. (1993). Records of pseudoscorpions associated with bats. Australian Mammalogy 16: 39-40.55. Harvey, M.S. (1993). The systematics of the Hyidae (Pseudoscorpionida: Neobisioidea). Invertebrate Taxonomy 7: 1-32.1992
54. Harvey, M.S. (1992). The phylogeny and classification of the Pseudoscorpionida (Chelicerata: Arachnida). Invertebrate Taxonomy 6: 1373-1435.53. Harvey, M.S. (1992). A new genus of myrmecophilous Chernetidae from southern Australia (Pseudoscorpionida). Records of the Western Australian Museum 15: 763-775.52. Main, B.Y. and Harvey, M.S. (1992). Type material of spiders (Chelicerata: Araneae) collected on the Michaelsen and Hartmeyer Expedition of 1905 and lodged in the Western Australian Museum. Records of the Western Australian Museum 15: 703-712.51. Harvey, M.S. (1992). A new species of Symphytognatha Hickman (Araneae: Symphytognathidae) from Western Australia. Records of the Western Australian Museum 15: 685-689.50. Harvey, M.S. (1992). The Schizomida (Chelicerata) of Australia. Invertebrate Taxonomy 6: 77-129.1991
49. Harvey, M.S. (1991). The cavernicolous pseudoscorpions (Chelicerata: Pseudoscorpionida) of Cape Range, Western Australia. Records of the Western Australian Museum 15: 487-502.48. Harvey, M.S. (1991). Notes on the genera Parahya Beier and Stenohya Beier (Pseudoscorpionida: Neobisiidae). Bulletin of the British Arachnological Society 8: 288-292.47. Harvey, M.S. and Waldock, J.M. (1991). Ecological survey of Abydos-Woodstock Reserve, Western Australia. Arachnid and myriapod fauna. Records of the Western Australian Museum, Supplement 37: 126-133.46. Harvey, M.S. (1991). The Pseudoscorpionida and Schizomida of the Kimberley rainforests. In: McKenzie, N.L., Johnson, R.B. and Kendrick, P.J. (eds), Kimberley Rainforests: 265-268. Surrey Beatty and Sons: Sydney.